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Justification Justification of Red List Category This species has an extremely large range, and hence does not approach the thresholds for Vulnerable under the range size criterion (Extent of Occurrence 30% decline over ten years or three generations). The population size is very large, and hence does not approach the thresholds for Vulnerable under the population size criterion (10% in ten years or three generations, or with a specified population structure). For these reasons the species is evaluated as Least Concern. Population justification The global population is estimated to number c.800,000-900,000 individuals (Wetlands International 2015), while the population in Russia has been estimated at c.10,000-100,000 breeding pairs and c.1,000-10,000 wintering individuals (Brazil 2009). The European population is estimated at 37,500-45,500 pairs, which equates to 75,000-91,000 mature individuals (BirdLife International 2015).

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Trend justification The overall population trend is decreasing, although one population may be stable and another has unknown trends (Wetlands International 2015). In North America, the species has undergone a small, statistically insignificant increase, over the last 40 years (data from Breeding Bird Survey and/or Christmas Bird Count: Butcher and Niven 2007) Note, however, that these surveys cover less than 50% of the species's range in North America. In Europe, the population trend is unknown (BirdLife International 2015).

Ecology Behaviour This species is fully migratory (del Hoyo et al. It breeds from late-June onwards, usually in well-dispersed solitary pairs, although in some areas it may also form loose colonies (Madge and Burn 1988; del Hoyo et al. 1992; Kear 2005). Males gather in vast flocks between July and August and undertake moult migrations to favoured marine areas often more than a thousand miles from the breeding grounds (Snow and Perrins 1998; Johnsgard 1978). The autumn migration to winter quarters, by both males and females), peaks between August and October, with stragglers still leaving the breeding grounds during September (Snow and Perrins 1998). The timing of the return passage in the spring is determined by the ice conditions and the thawing of inland breeding waters (Snow and Perrins 1998). Non-breeding birds generally oversummer south of the breeding range or may remain on the wintering grounds (Madge and Burn 1988).

The species is highly gregarious when not breeding, migrating in groups of up to c.1,000 individuals ( and aggregating into enormous flocks during the moulting period (e.g. 100,000 individuals off the western coast of Greenland)) and during the winter (Johnsgard 1978; Madge and Burn 1988; Snow and Perrins 1998). Habitat Breeding The species breeds on dry Artic tundra near freshwater lakes, pools, bogs, marshes, streams and small rivers the coast or up to 50 km (rarely up to 100 km) inland (del Hoyo et al. 1992, Kear 2005; Madge and Burn 1988). It shows a preference for shallow fresh waters with emergent vegetation for initial brood rearing, afterwards moving to more saline waters where the young fledge (Snow and Perrins 1998, Kear 2005). Non-breeding The species generally moults in sheltered fjords and bays with high densities of benthic fauna (Kear 2005), and winters at sea on deep offshore waters close to the edge of sea ice or in coastal areas with shallow waters (del Hoyo et al. 1992, Kear 2005).

Diet Its diet consists predominantly of animal matter such as benthic molluscs, crustaceans, larval insects (del Hoyo et al. Caddisflies and midges), echinoderms (e.g. Sea urchins and sand dollars) and other marine invertebrates, although the seeds and the vegetative parts of tundra plants, sedges and aquatic plants may also be taken on the breeding grounds, and algae, eelgrass Zostera spp.

And Ruppia maritima may be taken at sea (Johnsgard 1978; del Hoyo et al. Breeding site The nest is a slight hollow on dry ground and is usually positioned near water, in the open, or under the cover of driftwood, grass hummocks or rocks (Flint et al. 1984; del Hoyo et al. Threats Levels of hunting in this species are poorly regulated in parts of the range and may be unsustainable in some areas. Two to five percent of the population has been harvested annually in Alaska and Canada in recent years (Powell et al. 2012), accounting for 16,000 individuals (Rothe et al.

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